Re: Leaf Opacity

I would check that the UVs from your exported model are correct. If you are having a mismatch from what appears in SpeedTree to what is shown in the engine with the texture applied it could mean that the UVs were not exported correctly or that the export settings are incorrect. You should not need any tiling as was previously mentioned.

Perhaps you could use another DCC tool to double-check the UV layout from SpeedTree, such as Blender.
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